From:  Recent advances in the modulatory effects of kefir on the gut microbiota

 Modulation of gut microbiota composition by kefir in human and animal models.

Kefir preparation methodSample typeFeeding dosage/administrationExperiment durationDNA extractionOrgan used for microbial analysisReference
Commercially available kefirProfessional female soccer players, aged 18–29 years
(n = 21)
200 mL of kefir daily, through oral intake28 daysDiaRex® Stool Genomic DNA Extraction Kit (Cat No.: SD-0323, Diagen, Ankara, Turkey)Fecal samplesÖneş et al. [25]
Microbial changes result
Akkermansia muciniphila, Bifidobacterium, SCFA-related genera (e.g., Roseburia)
↓ Proteobacteria
Traditional kefir produced with grains by Danem, Inc. (kefirdanem.com, Suleyman Demirel University Technopark, Isparta, Turkey)Women with PCOS, aged between 18 and 40 years
(n = 17)
250 mL/day of kefir, through oral administration8 weeksDiaRex® Stool Genomic DNA Extraction KitFecal samplesÇıtar Dazıroğluet et al. [27]
Microbial changes result
↑ Firmicutes, Actinobacteria, and Lactococcus-related taxa
↓ Bacteroidetes, Proteobacteria, and Holdemania
Lifeway Foods® KefirICU patients,
(> 18 years)
(n = 54)
60 mL, followed by 120 mL after 12 h, then 240 mL of kefir daily, through oral or nasogastric administration4 weeksQiagen’s DNeasy 96 PowerSoil Pro QIA- cube HT Kit
(QIAGEN, Germantown, MD, USA)
Fecal samplesGupta et al. [28]
Microbial changes result
↑ Bacilli, Lactobacillus spp. (L. plantarum, L. reuteri, L. rhamnosus), Parvimonas and Dialister
Bifidobacterium longum
Kefir milk product (provided by Nourish Kefir)Healthy volunteers, aged from 18 to 65
(n = 9)
247 mL/day of kefir, through oral intake28 daysQIAamp DNA Stool Mini Kit (QIAGEN, UK)Fecal samplesWalsh et al. [26]
Microbial changes result
Lactococcus raffinolactis
Commercial kefir (the brand is not mentioned)Angora cats, age: 3.3 ± 2.5 years old
(n = 7; male: 5, female: 2)
30 mL/kg/day of kefir, orally administered14 daysSelective media for total mesophilic aerobic bacteria, coliform bacteria, Enterobacteriaceae, Enterococcus spp., Lactobacillus spp., Lactococcus spp., and yeast)Fecal samplesKabakçi et al. [31]
Microbial changes result
↑ Total mesophilic aerobic bacteria, Lactobacillus spp., Lactococcus spp., and yeast
Enterococcus spp.
Kefir grains were cultured in Irish whole full-fat cow’s milkMale BTBR T+ Itpr3tf/J mice, 5–6 months of age
(n = 9)
0.2 mL/day of kefir, through oral administration3 weeksQiaAmp PowerFecal Pro DNA KitCecal contentsvan de Wouw et al.
[29]
Microbial changes result
Lachnospiraceae bacterium A2
↓ Clostridiaceae and Clostridium
Kefir grains were independently cultured in Irish whole full-fat cow’s milk to prepare two distinct kefir types, Fr1 and UK4Male C57BL/6J mice, 8 weeks of age
(n = 48)
0.2 mL/day, through oral gavage3 weeksUsing QIAamp PowerFaecal DNA Kitileal, caecal, and faecal samplesvan de Wouw et al. [30]
Microbial changes result
Both kefir groups:
Lactobacillus reuteri (caecum and faeces), Eubacterium plexicaudatum (faeces and caecum), Bifidobacterium pseudolongum (ileum and caecum)
Lachnospiraceae bacterium 3_1_46FAA (caecum), Propionibacterium acnes (faeces), Bacillus amyloliquefaciens (faeces)
For Fr1 kefir group:
Parabacteroides goldsteinii (caecum), Bacteroides intestinalis (faeces), Anaerotruncus spp. (faeces), Parabacteroides goldsteinii (faeces)
For the UK4 kefir group:
Alistipes spp. (caecum)
Candidatus Arthromitus spp. (ileum)
Kefir peptides (KPs), fermented goat milk with traditional kefir grainsFemale C57BL/6J mice (ovariectomized model, simulating estrogen deficiency)
(n = 6)
0.1 mL/kg daily of KPs, through oral administration8 weeksQIAamp PowerFecal DNA Kit (Qiagen, Redwood, CA, USA)Cecal contentsTu et al. [33]
Microbial changes result
↑ Alloprevotella, Anaerostipes, Parasutterella, Romboutsia, Ruminococcus, and Streptococcus
Sterilized milk was fermented with kefir grainsHealthy adult dogs, 5.17 ± 2.32 years old
(n = 6; male: 4, female: 2)
200 mL of kefir daily, through oral administration2 weeksNucliSENS easyMAG instrument (bioMérieux, Marcy l’Etoile, France)Fecal samplesKim et al. [32]
Microbial changes result
↑ Prevotellaceae, Selenomonadaceae, Sutterellaceae, Catenibacterium mitsuokai, and LAB.
↓ Clostridiaceae, Fusobacteriaceae, Ruminococcaceae, Bacteroides uniformis, Fusobacterium mortiferum, Pseudoflavonifractor capillosus, and Fusicatenibacter saccharivorans
Kefir was prepared using the culture of DC1500I (Danisco, Olsztyn, Poland)Adults with metabolic syndrome, aged 18–65 years
(n = 12)
180 mL/day of kefir, consumed orally12 weeksQiagen Stool Mini Kit (Qiagen, Hilden, Germany)Fecal samplesBellikci-Koyu et al. [24]
Microbial changes result
↑ Firmicutes, Verrucomicrobia, Actinobacteria, Bacteroides, Clostridia, Lactobacillales, Bifidobacterium, and the Firmicutes/Bacteroidetes ratio
↓ Bacteroidetes, Proteobacteria, Prevotellaceae, Alistipes, and Veillonellaceae
Cow milk fermented with kefir grains vs. kefir starter cultureMale BALB/c mice
(n = 30)
0.3 mL/day of kefir, through oral gavage15 daysSelective culture media for Lactobacilli, LAB, lactic streptococci, yeasts/fungi, Bifidobacterium spp., and EnterobacteriaFecal samplesErdogan et al. [34]
Microbial changes result
Kefir grains group:
Lactobacillus spp., Streptococci, Lactobacillus acidophilus, Bifidobacterium spp., and yeast
Kefir starter culture group:
Lactobacillus spp., Streptococci, Lactobacillus acidophilus, and Bifidobacterium spp.

↑: increase; ↓: decrease; ICU: intensive care units; LAB: lactic acid bacteria; PCOS: polycystic ovary syndrome; SCFA: short-chain fatty acid; UK4 and Fr1: two types of kefirs made from different kefir grain communities.