From:  Fisetin in malignant nervous system tumors: molecular insights and translational advances

 FIS mechanism of action and biological effects.

MechanismFIS’s actionBiological effectTumor types affectedReference
Cell cycle modulationInduces G1/S arrestModifies cell cycle distributionGlioblastoma (U-138 MG)[45]
Apoptosis inductionActivates caspases-3, -8, and -9Increases apoptosis in glioblastoma cells
Nanoparticle encapsulationEncapsulated in PLGA nanoparticles (FIS-PLGA-NP4)Enhances stability and drug loading, cytotoxicity improvement
RadiostabilityExposure to ionizing radiation (25 kGy)Maintains structural integrity, suitable for sterilization
Combination therapySynergistic effect with chemotherapeuticsEnhances efficacy and reduces resistance
DNA damageInduces DNA strand breaks, measured by alkaline comet and γH2AX assayInitiates genotoxicity and the DNA damage response via p53 signalingGlioblastoma[46]
Apoptosis inductionInduces apoptosis through flow cytometry analysisIncreases cell death through apoptosis, thereby lowering overall cell survival
Senescence reductionReduces senescence induced by TMZDecreases the number of senescent cells, thereby increasing apoptosis
DNA repair inhibitionInhibits DNA repair pathways such as BERIncreases DNA damage, thereby increasing cell death
Enhancement of chemotherapyEnhances the genotoxic effect of TMZIncreases the efficiency of temozolomide by enhancing cell death and inhibiting senescence
Cell proliferation inhibitionInhibits cell proliferation in T98G cellsInhibits cell viability and growth in MTT assays as indicated by a lower IC50 in T98G cellsGlioblastoma (T98G cells)[47]
Apoptosis activationInduces apoptosis in T98G cells via caspases 3, 8, 9, and BaxEnhances apoptotic indices by assessing caspase activation, mitochondrial destabilisation, and DNA fragmentation
Cytotoxicity comparisonMore cytotoxic in T98G cells than carmustine (positive control)Causes cell death and cytotoxicity at lower concentrations in T98G cells than carmustine
Gene expression modulationIncreases CASPASE-3, CASPASE-9, BAX, and decreases BCL-2Regulates the expression of genes involved in apoptosis, thereby promoting apoptosis
Cell migration inhibitionSuppresses glioma cell migration at non-cytotoxic concentrationsReduces the migration and motility ability of malignant cellsGlioma (GBM8401 cells)[48]
Cell invasion suppressionInhibits invasion through the extracellular matrixInhibits the limits of the invasion potential without causing cytotoxicity
ADAM9 downregulationDecreases ADAM9 protein and mRNA expressionInsulates glioma development and invasion-related proteins
ERK1/2 signaling modulationInduces sustained phosphorylation of ERK1/2ERK1/2 is activated, leading to the inhibition of the ADAM9 expression
ERK1/2-dependent anti-invasive effectERK1/2 inhibition (U0126 or siERK) reverses FIS’s effectsElevates the ERK1/2 as a requisite mediator of the anti-invasive effect of FIS
Tumor microenvironment modulationAlters the tumor microenvironment, affects autophagy and apoptosis pathwaysEnhances a better treatment response and reduces treatment resistanceGlioblastoma, glioma[49]
Active targetingModification with cRGD peptidePenetrates the blood-brain barrier to reach glioma cellsGlioblastoma, glioma GL261 or U87-MG cells[56]
Redox-responsive releaseDisulfide bond-containing nanoparticlesBreaks down under the condition of increased glutathione, thus, liberating FIS and doxorubicin
Enhancement of drug uptakeFIS inhibits P-glycoprotein (P-gp)Modifies the cellular localization of doxorubicin and other chemotherapeutic agents
Cell cycle arrestInduction of G2/M phase arrestPrevents the multiplication and movement of glioma cells
Apoptosis promotionActivation of the caspase cascadeCauses oligonucleotide apoptosis via exogenous and endogenous mechanisms
Synergistic effects with DOXCombination with DOX-loaded nanoparticlesA synergistic increase in tumor suppression and a decrease in drug resistance
Cell proliferationDecreases cell proliferation in both TMZ-sensitive and TMZ-resistant cellsReduced cell viability and growth inhibitionGlioblastoma (TMZ-resistant T98G23 and TMZ-sensitive A172 cells)[57]
Cell migrationInhibits migration in TMZ-treated GB cells, enhancing the effects of TMZSuppression of the migratory behavior of glioblastoma cells
Combination therapyTMZ + FIS combination treatmentEnhanced chemotherapeutic efficacy, overcoming resistance in glioblastoma models
PI3K/Akt/CREB pathwayActivates PI3K/Akt/CREB signalingPromotes neuronal survival and neuroprotectionNeurodegenerative disorders[58]

FIS: fisetin; PLGA: poly(lactic-co-glycolic acid); NP: nanoparticle; γH2AX: gamma H2A histone family member X; TMZ: temozolomide; IC50: half-maximal inhibitory concentration; ADAM9: A disintegrin and metalloproteinase 9; ERK: extracellular signal-regulated kinase; BER: base excision repair; MTT: methylthiazolyldiphenyl-tetrazolium bromide; cRGD: cyclic arginine-glycine-aspartic acid (peptide); DOX: doxorubicin; PI3K: phosphatidylinositol 3-kinase; Akt: protein kinase B; mTOR: mechanistic (mammalian) target of rapamycin. CREB: cAMP response element-binding protein.