From:  Glucocorticoid receptor alpha as a core survival receptor: mechanisms, and implications for health and critical illness

 Comparative roles of GRα and MR across physiological systems and disease states.

Physiological systemGRα—key functionsMR—key functionsPathogenic imbalance/Clinical consequence
Systemic integrationMaster regulator of whole-body homeostasis; coordinates stress adaptation, metabolism, and mitochondrial energy production [58, 83, 84].Maintains electrolyte and volume balance; sustains baseline vascular tone [16, 86].↓ GRα → systemic inflammation, mitochondrial failure, and cardiovascular collapse [83]; ↑ MR → hypertension, vascular inflammation, fibrosis [16].
Immune systemRepresses excessive inflammation, promotes resolution, restores immune tolerance [83, 85].Supports macrophage survival and innate immune activation [85, 87].↓ GRα + ↑ MR → persistent inflammation, cytokine excess, impaired resolution [83, 85].
Cardiovascular/renalPreserves endothelial integrity via eNOS activation and anti-inflammatory signaling [83, 86].Regulates sodium retention, blood pressure, and vascular reactivity [16, 86].↓ GRα → endothelial dysfunction and vascular leakage [83]; ↑ MR → fibrosis, oxidative stress, and cardiometabolic injury [16].
Metabolic/hepaticEnhances gluconeogenesis under stress; maintains glucose and lipid balance [83, 84].Modulates insulin sensitivity and adipose inflammation [16].↓ GRα → energy failure and hypometabolism [83]; ↑ MR → adipose inflammation and insulin resistance [16].
CNS/neuroendocrineFacilitates stress adaptation, neuroplasticity, and emotional regulation [145, 146].Maintains basal excitability, circadian tone, and neuroendocrine stability [146].↑ GRα or ↓ MR → neuroinflammation, oxidative stress, excitotoxicity [145, 146].
Developmental/survivalRequired for postnatal viability; integrates transcriptional and non-genomic responses essential for stress tolerance [83, 145].Supports perinatal electrolyte and circulatory adaptation [86].Deletion of either → postnatal death (GRα = systemic failure; MR = salt-wasting collapse) [83, 86].

This table compares the complementary and opposing actions of GRα and MR across major physiological systems. GRα functions as the master integrator of whole-body homeostasis, coordinating metabolic, vascular, immune, and neuroendocrine responses required for stress adaptation and survival. MR primarily maintains electrolyte balance, vascular tone, and tissue-specific homeostasis. Loss of GRα results in multisystem failure, characterized by metabolic and immune collapse, whereas MR loss causes localized salt-wasting and vascular inflammation. Their imbalance underlies many critical-illness phenotypes, where GRα deficiency leads to systemic collapse and MR overactivation promotes inflammation and fibrosis. GRα: glucocorticoid receptor alpha; MR: mineralocorticoid receptor; eNOS: endothelial nitric oxide synthase; CNS: central nervous system. ↑: increased; ↓: decreased; →: leads to/results in.